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to double holliday junction which is shown here has its own remarkable properties and one of them which is illustrated is on the next slide here what happens is that this double holliday junction can actually be dissolved and and this is a property that cannot happen with a single Holliday Junction because it has no way to exchange strands in the way that Im talking about here the blooms helicase you remember that I mentioned blooms protein early on as being a defect of DNA repair well this protein turns out to be a helicase which can push on these two cross structures and push them towards each other when it does that this creates a steric problem because as you do this the DNA in between will become increasingly over-wound and and this would resist the action of the blooms Fela case but the blooms Gila case turns out to be associated with a topoisomerase which you probably know is an enzyme that can Nick one strand of DNA and rotate the DNA around the remaining strand taking up the