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In a holiday Junction, two homologous duplexes are linked by reciprocal exchange of DNA strands. Formation is initiated by a d-loop where one gray strand invades pink duplex, forming heteroduplex DNA regions. Cleavage and ligation create the Junction, with planar unfolded structure interacting with recombination proteins. Both duplexes have homo and heteroduplex regions, with four DNA strands involved. The Junction has two axes of symmetry, horizontal and vertical.
