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in a holiday Junction two homologous duplexes are linked by the reciprocal exchange of a pair of DNA strands the formation of a holiday Junction is initiated by the formation of a d-loop between homologous duplexes in which a strand from one duplex shown here in Gray invades the other duplex shown in pink regions of heteroduplex DNA are formed in which complementary strands from the two different duplexes are paired cleavage of the d-loop and subsequent ligation results in a holiday Junction which links the two duplexes holiday Junctions adopt a planar unfolded structure when they interact with recombination proteins notice that both of the linked duplexes contain regions of homo duplex where graystrand pairs with gray Strand and pink strand pairs with pink Strand and heteroduplex where great strand pairs with pink strand also notice that the holiday Junction involves four DNA strands there are two axes of symmetry in the holiday Junction one horizontal and one vertical strands of the
