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In this lecture, payroll deductions are defined as amounts withheld from an employee's gross pay, also known as withholdings. The key focus is on payroll taxes, which include federal income tax and FICA taxes for Social Security and Medicare. Additionally, deductions may include amounts withheld for benefits such as retirement and insurance plans. The process involves starting with gross pay, the actual earnings of the employee, from which various deductions, including federal taxes, are subtracted. This results in the net pay that the employee ultimately receives.