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the amino acid glutamate is the principal excitatory neurotransmitter in the brain and it is estimated that over half of all brain synapses release this agent like all amino acids glutamate has both a carboxyl and an amino functional group attached to the same carbon also attached to this carbon as a side chain glutamate is a non-essential amino acid that does not cross the blood-brain barrier and must therefore be synthesized in neurons from local precursors the most prevalent precursor for glutamate synthesis is glutamine which is released by glial cells once released glutamine is taken up into presynaptic terminals and metabolized to glutamate by the mitochondrial enzyme Glu Tammanys glutamate can also be synthesized by transamination of tuoc so glue to rate an intermediate as the tricarboxylic acid cycle hence some of the glucose metabolized by neurons can also be used for glutamate synthesis glutamate is packaged into synaptic vesicles by transporters termed Z glued at least thr