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To achieve clean and aesthetically pleasing URLs in PHP, traditional URLs (e.g., `profile.php?user=John&a=21`) can be transformed into a simpler format (e.g., `profile/John/21`). This is accomplished by modifying how the server interprets URLs through the `.htaccess` file. This file allows redirection of requests, enabling the server to handle requests without explicitly specifying the PHP file or query parameters. The tutorial suggests creating a new file and implementing specific code to achieve this redirection functionality, though it does not elaborate further on the technical details of the code.