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The speaker discusses how to customize the layout of a glyph in a text by anchoring it to the text and rasterizing the content. They explain how to ensure the background is transparent, select a custom layout, and adjust the alignment and spacing of the glyph relative to the text size. By setting the glyph to push the text down by 500 pixels, they show how to effectively position the glyph on the page. The speaker demonstrates how these export options affect an ePub file by resizing the window to show the changes in alignment and spacing of the glyph relative to the text.