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In this tutorial, the speaker discusses the phenomenon of light and dark adaptation, which refers to how our eyes adjust when transitioning between different lighting conditions. They illustrate this with relatable examples, such as leaving a dark movie theater and stepping into bright daylight, or moving from a brightly lit room to a dark space, like a basement. The speaker emphasizes the importance of understanding how our eyes continuously and dynamically adjust to varying light levels, setting the stage for a deeper exploration of the structures involved in this adaptation process.