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What springs to mind when we hear the words trait and inheritance? Many of us picture Mendel and his pea experiments. Mendels theory of inheritance states that organisms inherit two alleles for each trait, one from each parent. You could say its easy peasy. I apologise in advance for many terrible puns to come. Mendels theory works well with qualitative traits, where offspring can be placed into categories. For example, a cross between a homozygous smooth and homozygous wrinkled pea plant will produce a heterozygous pea that appears smooth as the smooth allele is dominant. However, not all traits can be explained so simply. Many traits are quantitative and display continuous variation, such as human height. How can we begin to make sense of these complex traits? In 1909, Nilsson-Ehle and East found that these quantitative traits result from multiple genes and their interaction with the environment. This discovery became the root of many pioneering experiments throughout the 20t
